Graduate 2019-2020

College of Natural and Health Sciences

School of Biological Sciences

BIO 501 Current Issues in Biology

Course varies in content, reviewing current issues of biology. S/U graded.
1- 6

Class Restriction

Exclude Freshman and Sophomore

Repeatable Status

Course is repeatable with a maximum of 6 credit hours

BIO 502 Biomedicine and Careers

This course will assist students in preparing for careers in biomedicine. Topics include biomedical ethics, interpersonal skill development, professionalism, the application process, interviewing skills and strategies to finance continued education.
1

Program Restrictions

Include Biomedical Science-MBS and Biomedical Science-MBS and BioSci:Non-Thesis-MS and BioSci:Non-Thesis-MS

Class Restriction

Exclude Freshman and Sophomore

BIO 508 Workshop

A variety of workshops on special topics within the discipline. Goals and objectives will emphasize the acquisition of general knowledge and skills in the discipline.
1- 3

Class Restriction

Exclude Freshman and Sophomore

Course Attribute

Variable Title Course

Repeatable Status

ST - Repeatable under different subtitles and course is repeatable with a maximum of 6 credit hours

BIO 511 Field Techniques in Biology

(1 Lecture, 1 Laboratory) The most common field techniques for both plant and animal biology, focusing on biodiversity.
2

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Course Fee Required

BIO 512 TA Development Seminar

Explores instructional strategies related to effective teaching of undergraduate laboratory learning environments. Intended to help develop teaching skills of graduate teaching assistants in the Biology program. S/U graded.
1

Class Restriction

Exclude Freshman and Sophomore

BIO 513 Professional Renewal

Update skills and knowledge of professionals in the discipline. Goals and objectives will be specifically directed at individual professional enhancement rather than the acquisition of general discipline knowledge or methodologies. S/U or letter graded.
1- 3

Class Restriction

Exclude Freshman and Sophomore

Course Attribute

Variable Title Course

Repeatable Status

ST - Repeatable under different subtitles and course is repeatable with a maximum of 6 credit hours

BIO 521 Advanced Genetics

Survey experimental advances in gene regulation of growth, maintenance and protection of all organisms including the human species with special emphasis on recent developments in the field.
3

Prerequisites

BIO 220: with minimum grade of C and CHEM 331: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 524 Genomics

Fundamental concepts of genomics. Introduction to the tools and techniques in the field of genomics, including genetic transformation, DNA sequencing, cDNA library synthesis, microarray, bioinformatics and sequencing of model organisms.
3

Class Restriction

Exclude Freshman and Sophomore

BIO 525 Molecular Genetics

Examination of protein synthesis, DNA replication, gene expression, gene structure, and regulation of gene expression.
3

Prerequisites

BIO 220: with minimum grade of C and (CHEM 381: with minimum grade of C or CHEM 481: with minimum grade of C)

Level Restriction

Include GR

Course Fee

Course Fee Required

BIO 526 Population Genetics

Investigation of empirical and theoretical population genetics. Topics will include: mutation, selection, migration, drift, genomics, quantitative genetics, human genetics, gene families, linkage disequilibrium, coalescence, and molecular clocks.
3

Prerequisites

BIO 111: with minimum grade of C and BIO 220: with minimum grade of C and MATH 131: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 527 Bioinformatics

An Introduction to the information technology of genomics and proteomics and their application to biological fields as diverse as evolution and medicine.
3

Prerequisites

BIO 111: with minimum grade of C and BIO 220: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 528 Developmental Biology

Exploration of animal development. Emphasis on the genetic basis of cell organization and identity during embryogenesis and differentiation. Topics include fertilization, gastrulation, axis specification, patterning, organogenesis, stem cells.
3

Prerequisites

BIO 220: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 529 Developmental Biology Lab

(3 laboratory) Classic model organisms such as sea urchin, fruit fly, chick, zebrafish, flat worm, and mouse will be used to observe and manipulate the events of animal development in living organisms.
1

Corequisites

BIO 528

Level Restriction

Include GR

Course Fee

Course Fee Required

BIO 530 Plant Systematics

(3 lecture, 3 laboratory) History of plant taxonomy, phylogenetic systematics, family recognition, and identification of local flora using keys. Native plant collection and field trips required.
4

Prerequisites

BIO 111: with minimum grade of C

Level Restriction

Include GR

Course Fee

Course Fee Required and Participation Fee Required

BIO 533 General Parasitology

(3 lecture, 3 laboratory) Study of the symbiotic relationships of parasitism as exemplified by typical parasites of humans, domesticated and wild animals, stressing life cycles, pathogenesis, systematics and hostparasite relationships.
4

Prerequisites

BIO 111: with minimum grade of C

Level Restriction

Include GR

BIO 534 Mammalogy

(3 lecture, 3 laboratory) Study of the functional biology, ecology and behaviors of mammals. Laboratory stresses the identification and ecology of Colorado species. Field work required.
4

Prerequisites

BIO 111: with minimum grade of C

Level Restriction

Include GR

Course Fee

Course Fee Required and Participation Fee Required

BIO 535 Survey of Fishes, Amphibians and Reptiles

(3 lecture, 3 laboratory) Investigation of the biology of the cold-blooded vertebrates including their evolution, ecology and behavior. Laboratory stresses the identification and biology of Colorado species.
4

Prerequisites

BIO 111: with minimum grade of C

Level Restriction

Include GR

Course Fee

Participation Fee Required

BIO 537 Morphogenesis of Algae and Fungi

(3 lecture, 3 laboratory) Study structure, morphogenesis and Phylogenetic relationships of organisms grouped with algae and fungi. Comparative study to explore multiple kingdom concepts. Ecological, medical and economic aspects of mycology and phycology explored.
4

Prerequisites

BIO 111: with minimum grade of C

Level Restriction

Include GR

Course Fee

Course Fee Required and Participation Fee Required

BIO 538 Ornithology

(3 lecture, 3 laboratory) Study of the evolution, general biology, diversity, ecology and behaviors of birds. Laboratory emphasis will be on identification.
4

Prerequisites

BIO 111: with minimum grade of C

Level Restriction

Include GR

BIO 539 Molecular Ecology

Investigation of the usage of molecular data to answer ecological questions. Topics include: marker selection, genetic characteristics of organisms, population analyses, phylogenetics conservation genetics, and phylogeography.
3

Prerequisites

BIO 220: with minimum grade of C

Corequisites

BIO 539L

Class Restriction

Exclude Freshman and Sophomore

BIO 539L Molecular Ecology Lab

(3 Laboratory) Laboratory to accompany BIO 539. Study of the techniques used to collect molecular data for ecological and evolutionary studies.
1

Prerequisites

BIO 220: with minimum grade of C

Corequisites

BIO 539

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Course Fee Required

BIO 540 Anatomy and Morphogenesis of Plants

(3 lecture, 3 laboratory) Study of the evolution of vascular plants as revealed by anatomy and morphology.
4

Prerequisites

BIO 111: with minimum grade of C and BIO 330: with minimum grade of C

Level Restriction

Include GR

BIO 541 Advanced Human Anatomy

(1 lecture, 6 laboratory) An in depth regional study of the anatomical structures of the human body. Spatial relationships among structures are emphasized. Laboratory includes organ and virtual dissection.
3

Prerequisites

BIO 341: with minimum grade of C and BIO 350: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Course Fee Required

BIO 542 Molecular and Cellular Laboratory

(2 Laboratory) Study of the theory and techniques currently used to investigate cells and molecules. Development of the laboratory and problem solving skills to successfully conduct experiments.
2

Prerequisites

BIO 220: with minimum grade of C and (May concurrently take BIO 525: with minimum grade of C or May concurrently take BIO 550: with minimum grade of C)

Level Restriction

Include GR

Course Fee

Course Fee Required

BIO 544 Comparative Vertebrate Adaptations

(3 lecture, 3 laboratory) Study of the diverse adaptations of vertebrates, from morphological to molecular levels of organization; includes significant overview of relevant primary literature.
4

Prerequisites

BIO 345: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Participation Fee Required

BIO 545 Pathogenic Microbiology

This lecture course addresses medically-relevant microorganisms and disease. Particular emphasis is placed upon mechanisms of pathogenesis, including microbial virulence factors and immunopathology, epidemiology and public health.
3

Prerequisites

BIO 351: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 546 Pathogenic Microbiology Laboratory

Students will learn techniques for the isolation, propagation and identification of pathogenic microorganisms.
1

Prerequisites

BIO 351: with minimum grade of C

Corequisites

BIO 545

Class Restriction

Exclude Freshman and Sophomore

BIO 547 Immunology and Disease

Explore the cause of disease in animals with a detailed emphasis on functions of the immune system that provide resistance to disease.
3

Prerequisites

BIO 220: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 548 Virology

Study of the principles of virology, including biochemistry, structure, taxonomy, transmission, isolation and identification strategies.
3

Prerequisites

BIO 351: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 549 Virology Laboratory

(3 laboratory) Study and development of laboratory skills in animal virology, including virus propagation, quantification, and detection assays.
1

Corequisites

BIO 548

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Course Fee Required

BIO 550 Cell Physiology

Study of the structural and functional relationships among biological macromolecules, cell organelles and cellular processes.
3

Prerequisites

BIO 220: with minimum grade of C and (CHEM 231: with minimum grade of C or CHEM 331: with minimum grade of C)

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Course Fee Required

BIO 552 Advanced Systemic Physiology: Homeostats

Advanced systemic physiology, with an emphasis on humans. Essential and advanced topics in systemic physiology, focusing on homeostatic systems.
3

Prerequisites

BIO 210: with minimum grade of C and (CHEM 381: with minimum grade of C or CHEM 481: with minimum grade of C)

Class Restriction

Exclude Freshman and Sophomore

BIO 553 Advanced Systemic Physiology: Control Systems

In-depth examination of the physiological control systems in the human.
3

Prerequisites

BIO 210: with minimum grade of C and (CHEM 381: with minimum grade of C or CHEM 481: with minimum grade of C)

Class Restriction

Exclude Freshman and Sophomore

BIO 554 General Plant Physiology

(3 lecture, 3 laboratory) Study physiological factors influencing the chemical and structural composition of plant absorption and utilization of water and minerals; photosynthesis, translocation, respiration, nitrogen metabolism and growth and development.
4

Prerequisites

BIO 111 and (CHEM 231 or CHEM 281 and CHEM 281L or CHEM 331 and CHEM 331L)

Level Restriction

Include GR

Course Fee

Course Fee Required

BIO 555 Reproductive Biology

Examination of the anatomy, physiology and mechanisms of reproduction with an emphasis on mammals.
3

Prerequisites

BIO 110: with minimum grade of C and CHEM 331: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 556 Current Topics in Biomedical Research

Examine current biomedical research, molecular, cellular, and physiological studies of human and model systems. Mechanisms regulating normal function, effects of toxicants and drugs, and the role of genetic mutations.
2

Prerequisites

BIO 110: with minimum grade of C and CHEM 331: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

Repeatable Status

Course is repeatable with no limitations

BIO 557 Mammalian Physiology I Laboratory

(3 laboratory)Laboratory experimentation, small group discussion and active learning exercises to complement BIO 552.
1

Corequisites

BIO 552

Class Restriction

Exclude Freshman and Sophomore

BIO 558 Mammalian Physiology II Laboratory

(3 laboratory) Laboratory experimentation, small group discussion and active learning exercises to compliment BIO 553.
1

Corequisites

BIO 553

Class Restriction

Exclude Freshman and Sophomore

BIO 559 Plant Reproductive Growth and Development

Advanced study of the genetic, physiological and environmental factors that influence reproductive development in flowering plants.
3

Prerequisites

BIO 354: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 560 Behavioral Ecology

Study of the principles of behavioral ecology, including the evolutionary basis of behavior, economic models of behavior, interspecific interactions, and social behavior.
3

Prerequisites

BIO 362: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Participation Fee Required

BIO 561 Plant Ecology

Examination of plant communities, effects of environmental factors on plants, as well as community measurement and evaluation, modeling techniques, energy flow through ecosystems, and community dynamics.
3

Prerequisites

BIO 110: with minimum grade of C and BIO 360: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Participation Fee Required

BIO 562 Conservation Biology

Examination of the fundamental principles of conservation biology (biodiversity, habitat degradation, extinction, restoration, planning) with discussion of current topics (climate change, conservation genetics, landscape vs. ecosystem conservation, sustainable development).
3

Prerequisites

BIO 111: with minimum grade of C

Level Restriction

Include GR

BIO 563 Animal Communication

Study of the principles of animal communication, including signal modalities, signal design and signal evolution.
3

Prerequisites

BIO 362: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

Course Fee

Participation Fee Required

BIO 564 Plant Ecology Lab

Examines methods of plant ecology including monitoring community structure and function, population dynamics, physiological tolerance, and date analysis. The lab will include both field and laboratory experiments.
1

Prerequisites

BIO 110: with minimum grade of C and BIO 360: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 565 Current Topics in Ecology and Evolutionary Research

Examine topics in current ecology and evolutionary biology research. Focus on speciation, microevolution, range shifts, species interactions, and ecosystems. Topics will include both applied and basic studies.
2

Prerequisites

BIO 111: with minimum grade of C and BIO 360: with minimum grade of C and BIO 465: with minimum grade of C

Degree Restriction

Include MA and MS and MBS and MACC and MAT and MBA and MPH

Class Restriction

Exclude Freshman and Sophomore

Repeatable Status

Course is repeatable with a maximum of 6 credit hours

BIO 566 Animal Physiological Ecology

Study of animal function and how environmental conditions influence the physiology of animals. Emphasis will be on vertebrate systems but invertebrates will be included.
3

Prerequisites

BIO 110: with minimum grade of C and BIO 360: with minimum grade of C

Level Restriction

Include GR

BIO 567 Animal Physiological Ecology Laboratory

(3 laboratory) Laboratory to accompany BIO 566 - animal physiological ecology. Includes instruction on the theory behind and use of physiological equipment/instruments and an examination of how environmental conditions affect animal function.
1

Prerequisites

BIO 110: with minimum grade of C and BIO 360: with minimum grade of C and May concurrently take BIO 566: with minimum grade of C

Level Restriction

Include GR

BIO 568 Disturbance Ecology

Study of natural and anthropogenic disturbances (fire, flood, insects outbreaks) effects on populations, communities and ecosystem function.
3

Prerequisites

BIO 110: with minimum grade of C and BIO 360: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 569 Conceptual Issues in Evolution

Study of the guiding principles in modern evolutionary biology. Focus on concepts of fitness, selection, adaptation, form and function, concerted evolution and the nature of biological variation.
2

Prerequisites

BIO 465: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 570 Coevolution in Ecological Systems

Survey of contemporary primary literature on coevolution and ecology, including symbiosis, mutualism, competition, pollination, mimicry, parasitism, seed dispersal, etc.
3

Prerequisites

BIO 360: with minimum grade of C and BIO 465: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 572 Species and Speciation

Examination of the theoretical and pragmatic perspectives of species recognition and critical review of speciation. Includes considerations of species recognition in context of conservation biology.
3

Prerequisites

BIO 465: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 573 Animal Communication Laboratory

Laboratory to accompany BIO 563 Animal Communication. Study and application of research techniques in the field of Animal Communication. Students will plan and conduct research.
1

Prerequisites

BIO 362: with minimum grade of C

Corequisites

BIO 563

Class Restriction

Exclude Freshman and Sophomore

BIO 574 Behavioral Ecology Laboratory

Laboratory to accompany BIO 560 Behavioral Ecology. Study and application of research techniques in the field of Behavioral Ecology. Students will plan and conduct research.
1

Prerequisites

BIO 362: with minimum grade of C

Corequisites

BIO 560

Class Restriction

Exclude Freshman and Sophomore

BIO 576 Pharmacology

A mechanistic study of pharmacokinetics (absorption, distribution, metabolism and excretion of drugs) and pharmacodynamics (drug actions and interactions). Examples of important bioactive chemicals will be discussed.
3

Prerequisites

(BIO 246: with minimum grade of C or BIO 350: with minimum grade of C or BIO 552: with minimum grade of C)

Level Restriction

Include GR

Class Restriction

Exclude Freshman and Sophomore and Junior and Senior

BIO 578 Pathobiology

Causes and mechanisms of disease at the molecular, cellular, and tissue levels. Cellular homeostasis and adaptive mechanisms, histopathology, toxins/toxicants, infectious disease, senescence, and target tissues.
3

Prerequisites

(May concurrently take BIO 341: with minimum grade of C or May concurrently take BIO 541: with minimum grade of C) and (May concurrently take BIO 350: with minimum grade of C or May concurrently take BIO 552: with minimum grade of C or May concurrently take BIO 553: with minimum grade of C) and (May concurrently take BIO 381: with minimum grade of C or May concurrently take BIO 547: with minimum grade of C)

Class Restriction

Exclude Freshman and Sophomore

BIO 580 Mammalian Endocrinology

Survey of endocrinology including neuroendocrine physiology, mechanisms of action, regulation of homeostasis, endocrine dysfunction, quantifying hormones.
3

Prerequisites

BIO 210: with minimum grade of C and BIO 350: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 581 Neurobiology

Examination of the development, anatomy, and physiology of the nervous system, with an emphasis on humans. Topics will include mechanisms mediating behavior and major brain diseases.
3

Prerequisites

BIO 350: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 582 Cancer Biology

Study the principles of cancer biology including the mechanisms involved in its progression; the roles of DNA damage, heredity, and the environment; and treatment and prevention.
3

Prerequisites

BIO 220: with minimum grade of C

Class Restriction

Exclude Freshman and Sophomore

BIO 584 Current Topics in Biological Education Research

Examine current topics in biological education research. Focus on teaching, learning, and assessment of biology as well as school policy and curriculum development.
2

Level Restriction

Include GR

Repeatable Status

Course is repeatable with a maximum of 6 credit hours

BIO 585 Topics in Field Biology

After preparatory work, study biology in the field on a state, national or international basis.
1-10

Class Restriction

Exclude Freshman and Sophomore

Course Attribute

Variable Title Course

Repeatable Status

Course is repeatable with a maximum of 10 credit hours

BIO 591 Biology Roundtable

An intensive facilitated discussion on current and cutting-edge topics in the Biological Sciences taken from peer reviewed literature and input from visiting scholars. S/U graded.
1

Class Restriction

Exclude Freshman and Sophomore

Repeatable Status

Course is repeatable with no limitations

BIO 592 Internship in Biological Sciences

On-the-job experience in professional areas under the supervision of an area specialist. S/U graded.
1- 3

Class Restriction

Exclude Freshman and Sophomore

BIO 594 Foundations of Biological Research

Introduction to the scientific method and experimental design, written and oral presentation skills, grantsmanship, ethics and responsible scientific conduct.
2

Class Restriction

Exclude Freshman and Sophomore

BIO 595 Special Topics in Biology

Study of any biological topic at an advanced level.
1- 6

Class Restriction

Exclude Freshman and Sophomore

Course Attribute

Variable Title Course

Repeatable Status

Course is repeatable with a maximum of 6 credit hours

BIO 596 Foundations of Clinical Research

This course examines the components of clinical research including design, conduct, analysis of trials, and dissemination of results.
2

Degree Restriction

Include MBS

Level Restriction

Include 00

Class Restriction

Include Master

BIO 622 Directed Studies

Individualized investigation under the direct supervision of a faculty member. (Minimum of 37.5 clock hours required per credit hour.)
1- 4

Level Restriction

Include GR

Course Attribute

Variable Title Course

Repeatable Status

ST - Repeatable under different subtitles with no limits on the number of times it can be repeated

BIO 631 Teaching College Biology

This course prepares students for everyday classroom teaching experiences in college biology including small and large classrooms and laboratory settings using a variety of best-practices teaching strategies.
3

Level Restriction

Include GR

BIO 632 College Biology Course Development

This course prepares students to effectively design college biology course curriculum using backwards design to develop learning objectives, syllabi, student assessments, and evaluation of course effectiveness.
3

Level Restriction

Include GR

BIO 691 Seminar in Biological Sciences

Invited speakers will present research topics in content biology and biology education. Graduate students will also present their research proposals and final defenses of their theses or dissertations.
1

Level Restriction

Include GR

Repeatable Status

Course is repeatable with no limitations

BIO 693 Review of Biological Literature

This course teaches students how to conduct a thorough review of scientific literature on a focused biological topic and prepare a written comprehensive report following standard scientific format and style.
3

Major/Minor Restriction

Include Biological Sciences

Type of Restriction For Major/Minor Code

Major(s)

Level Restriction

Include GR

BIO 697 Graduate Research

Qualified graduate students spend a minimum of 38 clock hours per semester hour on a research problem. Before credit is given, a well written report must be submitted to the instructor.
1- 6

Level Restriction

Include GR

Repeatable Status

Course is repeatable with a maximum of 6 credit hours

BIO 699 Thesis

Thesis S/U graded.
1- 6

Level Restriction

Include GR

Repeatable Status

Course is repeatable with no limitations

BIO 792 Doctoral Supervised Teaching

Experience supervised college level biological sciences teaching. Develop course outlines and effective teaching techniques. Students must have completed oral and written comprehensive exams prior to being eligible for this course.
1- 6

Level Restriction

Include GR

Repeatable Status

Course is repeatable with a maximum of 6 credit hours

BIO 797 Doctoral Proposal Research

Four credits required of all doctoral students before admission to candidacy. S/U graded.
1- 4

Level Restriction

Include GR

Repeatable Status

Course is repeatable with a maximum of 4 credit hours

BIO 799 Doctoral Dissertation

Twelve credits required of all doctoral candidates. S/U graded.
1-12

Level Restriction

Include GR

Repeatable Status

Course is repeatable with no limitations